It wont reset itself , it will slow retune it self to the AFR's that are in the template AFR table in the ECU.
Doing a manual reset will help this happen quickly, but from our knowledge on the 350Z tuning , you will have no issues running the car with the mods fitted before you get a remap, the car may feel a little sluggish mid range thou until you get the remap.
cheers
It wont be the plenum spacer making the car run lean, the HFC,s make the car runs lean before a remap.
Mark